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studying His creations, Haldane is said to have replied, “He seems to have an inordinate fondness for beetles.” Although the scope for describing new beetles and other small animals remains enormous (Fig. 3.2), much of our attention in cataloging biological diversity shifted toward even smaller creatures following a classic study undertaken by a group of Norwegian microbiologists (Torsvik et al. 1990a, b). From two tiny soil samples – 1 gram of forest soil and 1 gram of marine sediment – scientists extracted first the bacteria and then the bacteria’s DNA. They then estimated the diversity of DNA strands, made a conservative assumption that bacteria are different species if less than 70% of their DNA is identical, and arrived at a rough estimate that each sample contained over 4000 species of bacteria, with little or no overlap in species between the two samples. Finding over 4000 bacteria species in a pinch of Norwegian soil was doubly impressive when you realize that, in 1990, 4000 was roughly the number of species of bacteria that had ever been described from all environments in the entire world. Today the number of described bacteria species has risen to about 10,000. Estimates of undescribed species still range greatly, up into the millions (Haegeman et al. 2013), with sizable numbers coming from the microbiome – the enormous suite of microbes that occupy the bodies of humans and other species (Huttenhower et al. 2012).

Photos depict the depth of unexplored biodiversity is greatest among small species as exemplified by this amphipod (a), recently identified from the Antarctic Ocean, or this cave beetle (b) from China.

      ([a] Cédric d’Udekem d’Acoz/Royal Belgian Institute of Natural Sciences; [b] Sunbin Huang and Mingyi Tian)

Photos depict a few caterpillars representing ten sibling species of what was long thought to be a single butterfly species, Astraptes fulgerator.

      (Dan Janzen/National Academy of Sciences, U.S.A)

Photos depict a species' intrinsic value is independent of its relationship with any other species as depicted on the left, whereas its instrumental value depends on its importance to other species, including people.

      (Malcolm L. Hunter Jr., author)

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